Fan, Tianqu https://orcid.org/0009-0001-4152-472X
Zhao, Jiangsha
Li, Ling
Cui, Meihua
Zhang, Jiawei https://orcid.org/0000-0002-5782-5543
Chi, Tian https://orcid.org/0000-0002-9675-6990
Shi, Shuo https://orcid.org/0000-0003-1728-3601
Funding for this research was provided by:
National Natural Science Foundation of China (32570848)
Natural Science Foundation of Shanghai (25ZR1401254)
Article History
Received: 22 October 2025
Revised: 20 March 2026
Accepted: 9 April 2026
First Online: 22 April 2026
Competing interests
: The authors declare no competing interests.
: All methods were performed in accordance with the relevant guidelines and regulations. For the use of human clinical samples, ethical approval of the human tissue array was granted by the Ethics Committee of Shanghai Outdo Biotech Co., Ltd (Approval No. SHYJS-CP-1507006). Written informed consent was obtained from all participants. For animal experiments, the study was approved by the Institutional Animal Care and Use Committee (IACUC) of ShanghaiTech University (Approval No. 20210420001 and 20250919001).