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Authors
Mikawa, Takumi https://orcid.org/0000-0003-0980-2379
Kameda, Masahiro
Ikari, Sumiko
Shibata, Eri
Liu, Shuyu
Miyagawa, Sawa
Ono, Koh https://orcid.org/0000-0002-4163-980X
Ito, Tomiko
Yoshizawa, Akihiko
Sugimoto, Masataka
Shibuya, Shuichi
Shimizu, Takahiko
Almunia, Julio
Ogiso, Noboru
Revêchon, Gwladys https://orcid.org/0000-0002-4824-6793
Palazzo, Alberta
Bernard, David https://orcid.org/0000-0002-1557-2074
Kanda, Hiroaki
Soga, Tomoyoshi
Takubo, Keiyo
Morioka, Shin
Sasaki, Junko
Sasaki, Takehiko https://orcid.org/0000-0003-1837-3748
Itamoto, Akihiro
Fujii, Takayuki
Seno, Hiroshi
Inagaki, Nobuya
Kondoh, Hiroshi
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (26310103)
MEXT | Japan Society for the Promotion of Science (16K15412)
MEXT | Japan Society for the Promotion of Science (17H04150)
MEXT | Japan Society for the Promotion of Science (18K18451)
MEXT | Japan Society for the Promotion of Science (20K20658)
MEXT | Japan Society for the Promotion of Science (23K27507)
MEXT | Japan Society for the Promotion of Science (23H03884)
MEXT | Japan Society for the Promotion of Science (22K06221)
MEXT | Japan Society for the Promotion of Science (19K07887)
MEXT | Japan Society for the Promotion of Science (21K15661)
MEXT | Japan Society for the Promotion of Science (23K27595)
MEXT | Japan Society for the Promotion of Science (20H04116)
MEXT | Japan Society for the Promotion of Science (24K02871)
MEXT | Japan Society for the Promotion of Science (20H03583)
MEXT | Japan Society for the Promotion of Science (22K1960)
MEXT | Japan Society for the Promotion of Science (17H03574)
MEXT | Japan Society for the Promotion of Science (23K21640)
MEXT | Japan Society for the Promotion of Science (22H04922)
MEXT | Japan Society for the Promotion of Science (JP24H00640)
The GAP fund program, the Incubation Program and IPG-Advance of Kyoto University
Institut National Du Cancer (PLBio No. 2018-144)
Institut National Du Cancer (INCA TABAC 17-007)
MEXT | JST | Core Research for Evolutional Science and Technology (JP20gm1210011)
MEXT | JST | Core Research for Evolutional Science and Technology (JP25gm1710007)
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Article History
Received: 20 February 2025
Revised: 29 September 2025
Accepted: 11 November 2025
First Online: 15 December 2025
Competing interests
: Related international applications of patents include WO2023/038027 and WO2024/135719, which have been filed in multiple countries including Japan and the United States.
