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Authors
Ito, Akane
Usui, Noriyoshi https://orcid.org/0000-0001-8079-8013
Doi, Miyuki https://orcid.org/0009-0007-9847-1861
Shimada, Shoichi https://orcid.org/0000-0001-5369-1069
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (23H02837)
MEXT | Japan Society for the Promotion of Science (24K22230)
MEXT | Japan Society for the Promotion of Science (23K14443)
Uehara Memorial Foundation
Takeda Science Foundation
Naito Foundation
Mochida Memorial Foundation for Medical and Pharmaceutical Research
SENSHIN Medical Research Foundation
Osaka Medical Research Foundation for Intractable Diseases
Inamori Foundation
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Article History
Received: 17 December 2024
Revised: 30 July 2025
Accepted: 1 September 2025
First Online: 6 October 2025
Competing interests
: The authors declare no competing interests.
: All experimental procedures were conducted in accordance with the ARRIVE guidelines and relevant institutional regulations. Animal experiments were approved by the Animal Research Committee of The University of Osaka (Approval No. 27-010). This study did not involve human participants; therefore, informed consent was not applicable.
