Li, Jinbo https://orcid.org/0000-0003-1565-4173
Fan, Jinxiao
Yao, Zhenqiang https://orcid.org/0000-0002-4664-3648
Jiao, Han
Duan, Rong
Sanders, James O.
Mesfin, Addisu
Xing, Lianping https://orcid.org/0000-0002-2588-7766
Boyce, Brendan F. https://orcid.org/0000-0002-3587-4854
Funding for this research was provided by:
U.S. Department of Health & Human Services | NIH | National Institute of Arthritis and Musculoskeletal and Skin Diseases (R01AR43510)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (R01AG049994)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (R01AG049994)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (R01AG076731)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (R01AG059775)
National Natural Science Foundation of China (82472481)
National Natural Science Foundation of China (82172469)
National Natural Science Foundation of China (82172469)
National Natural Science Foundation of China (82472481)
Article History
Received: 19 November 2024
Revised: 16 January 2026
Accepted: 11 February 2026
First Online: 18 May 2026
Competing interests
: The authors declare no competing interests.