Article History
Received: 13 May 2025
Revised: 21 October 2025
Accepted: 12 November 2025
First Online: 24 November 2025
Competing interests
: The authors declare no competing interests.
: All animal experiments were approved by the Institutional Animal Care and Use Committee of University of Southern California (NO. 11673), and were conducted in accordance with the committee’s institutional guidelines.