Miao, Lingling
Collado, Loren
Barkdull, Savannah https://orcid.org/0009-0004-1162-3665
Hallaert, Patrick https://orcid.org/0009-0007-1998-1119
Martin, Mackenzie R.
Gryder, Berkley E.
Kelly, Michael C. https://orcid.org/0000-0003-0654-2778
Dell’Orso, Stefania
Kelley, Matthew W. https://orcid.org/0000-0001-7367-8697
Brownell, Isaac https://orcid.org/0000-0002-0090-9914
Funding for this research was provided by:
U.S. Department of Health & Human Services | NIH | National Institute of Arthritis and Musculoskeletal and Skin Diseases (Intramural Research Program: ZIA AR041221)
Article History
Received: 3 December 2024
Revised: 2 December 2025
Accepted: 21 January 2026
First Online: 3 February 2026
Competing interests
: The authors declare no competing interests.
: All animal experiments were carried out in accordance with institutional guidelines and IACUC-approved protocols at the National Cancer Institute and the National Institute of Arthritis and Musculoskeletal and Skin Diseases, NIH.