Ko, Jeong Yeon
Kweon, Tae Hyun
Jung, Hyeryeon https://orcid.org/0000-0001-5179-8604
Kang, Jingu
Kim, Yeolhoe https://orcid.org/0000-0003-2832-2493
Kim, Yun Ju
Shin, Donghyuk https://orcid.org/0000-0002-8272-6133
Yang, Seong Wook
Lee, Myeong Min
Hong, Jun Young
Lim, Jae-Min
Yi, Eugene C.
Cho, Jin Won
Yang, Won Ho https://orcid.org/0000-0001-6214-1421
Funding for this research was provided by:
National Research Foundation of Korea (NRF-RS-2023-00213643)
National Research Foundation of Korea (NRF-2018R1A6A1A03025607)
Article History
Received: 11 November 2024
Revised: 13 October 2025
Accepted: 3 November 2025
First Online: 9 January 2026
Competing interests
: The authors declare no competing interests.
: The authors utilized ChatGPT-4o to assist with grammar correction and language refinement during the preparation of this manuscript. All content was subsequently reviewed and revised by the authors, who take full responsibility for the final version of the publication.
: All experimental procedures were performed in accordance with the relevant guidelines and regulations. All animal experiments were approved by the Institutional Animal Care and Use Committees of the Laboratory Animal Research Center at Yonsei University (IACUC-A-20221018-41) and conducted in accordance with institutional and national guidelines for the care and use of laboratory animals.