Adachi, Takumi
Matsushita, Kazufumi
Yasuda, Koubun
Nakahira, Masakiyo
Imasaka, Mai
Sugimoto, Michihiko https://orcid.org/0000-0002-5888-5054
Yang, Zhuohao https://orcid.org/0009-0000-0983-7462
Kobiyama, Kouji
Hayashi, Tomoya
Ishii, Ken J. https://orcid.org/0000-0002-6728-3872
Yoshioka, Yasuo
Ohmuraya, Masaki
Shirasaki, Yoshitaka https://orcid.org/0000-0002-3808-1630
Kuroda, Etsushi https://orcid.org/0000-0002-7198-2190
Article History
Received: 14 July 2025
Revised: 3 November 2025
Accepted: 4 December 2025
First Online: 22 December 2025
Competing interests
: Yasuo Yoshioka was employed by the Research Foundation for Microbial Diseases at The University of Osaka.
: All animal experiments were performed according to the Guidelines for the Care and Use of Animals approved by Hyogo Medical University. (Approval numbers 22-035AG and 23-018AGP).