Ali, Mohamad Moustafa https://orcid.org/0000-0002-4902-0550
Itoh, Yuka
Badji, Aisha Mariama Pereira
Gallant, Sarah https://orcid.org/0009-0003-9742-4706
Tsirigoti, Chrysoula https://orcid.org/0000-0001-6554-738X
Bai, Yu https://orcid.org/0009-0002-0812-8917
Filipek-Górniok, Beata
Miyazawa, Keiji https://orcid.org/0000-0002-7835-7718
Heldin, Carl-Henrik https://orcid.org/0000-0002-9508-896X
Moustakas, Aristidis https://orcid.org/0000-0001-9131-3827
Article History
Received: 26 June 2025
Revised: 8 December 2025
Accepted: 21 January 2026
First Online: 4 February 2026
Competing interests
: The authors declare no competing interests.
: All methods were performed in accordance with the relevant guidelines and regulations. This study does not include human subjects. The zebrafish experiments were performed at the Genome Engineering Zebrafish National Facility (currently DanioReadout, SciLifeLab Uppsala University), Sweden, that holds the appropriate ethical approval from the Swedish Board of Agriculture (5.8.18-10590/2018 and 5.8.18-06282/2023).