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Authors
Aikawa, Shizu https://orcid.org/0000-0001-7407-4051
Shimizu-Hirota, Ryoko
Sakashita, Akihiko
He, Xueting https://orcid.org/0009-0003-6101-5046
Hiratsuka, Daiki https://orcid.org/0000-0001-7128-2859
Ishizawa, Chihiro
Iida, Rei
Fukui, Yamato
Hiraoka, Takehiro
Matsuo, Mitsunori
Takeda, Norihiko
Ikawa, Masahito https://orcid.org/0000-0001-9859-6217
Osuga, Yutaka
Hirota, Yasushi https://orcid.org/0000-0003-0241-9780
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (24K21911)
MEXT | Japan Society for the Promotion of Science (25H01065)
MEXT | Japan Society for the Promotion of Science (23K27176)
MEXT | Japan Society for the Promotion of Science (23K08278)
MEXT | Japan Society for the Promotion of Science (19K16022)
MEXT | Japan Society for the Promotion of Science (24K22157)
MEXT | Japan Society for the Promotion of Science (25K02779)
Japan Agency for Medical Research and Development (JP25gk0210039)
Japan Agency for Medical Research and Development (JP25gn0110097)
Japan Agency for Medical Research and Development (JP24gn0110069)
Japan Agency for Medical Research and Development (JP24lk0310083)
Japan Agency for Medical Research and Development (JP25gk0210042)
Japan Agency for Medical Research and Development (JP25gk0210045)
Japan Agency for Medical Research and Development (JP25gn0110085)
MEXT | Japan Science and Technology Agency (JPMJFR210H)
Astellas Foundation for Research on Metabolic Disorders (N/A)
Naito Foundation (N/A)
Inoue Foundation for Science (N/A)
Uehara Memorial Foundation (N/A)
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Article History
Received: 26 June 2025
Revised: 7 January 2026
Accepted: 10 February 2026
First Online: 9 March 2026
Competing interests
: The authors declare no competing interests.
: All methods were performed in accordance with the relevant guidelines and regulations. All animal experiments were approved by the Institutional Animal Experiment Committee of the University of Tokyo Graduate School of Medicine (Approval numbers P16-066, P20-076, and A2023M165).
