Xiao, Yuqi https://orcid.org/0009-0007-5576-7734
He, Mingyuan
Yao, Han
Song, Yimeng
Hu, Yu
Liu, Xinglong
Bai, Yang
Zhang, Jianghong https://orcid.org/0000-0003-4884-6411
Shao, Chunlin https://orcid.org/0000-0001-9336-9912
Funding for this research was provided by:
National Natural Science Foundation of China (12235004)
National Natural Science Foundation of China (12175044)
National Natural Science Foundation of China (12375338, 82173461)
Article History
Received: 26 November 2025
Revised: 1 April 2026
Accepted: 18 April 2026
First Online: 3 May 2026
Competing interests
: The authors declare no competing interests.
: This study complied with the Declaration of Helsinki and the Basel Declaration. Protocols involving human subjects and their data were reviewed and approved by the Ethics Committee of the China-Japan Union Hospital of Jilin University (2022-KYYS-048), with informed consent obtained from all participants. All animal procedures were approved by the Animal Welfare and Ethics Committee of Fudan University and conducted in strict accordance with relevant guidelines (2023-FSYX-05JZS).