Wei, Yang https://orcid.org/0009-0003-3709-2949
Sun, Dayan
Wu, Fei
Zhang, Shixuan
Cai, Bowen
Ma, Yanyun
Zheng, Hongxiang
Shi, Xiangguang https://orcid.org/0000-0002-3907-5822
Li, Yi
Le, Shiguan
Zhou, Xiang
Jin, Li https://orcid.org/0000-0001-9201-2321
Wang, Jiucun https://orcid.org/0000-0003-2765-0620
Funding for this research was provided by:
National Natural Science Foundation of China (U23A20475, 32288101, 82300574, 31871436)
Article History
Received: 7 August 2025
Revised: 21 November 2025
Accepted: 12 December 2025
First Online: 6 January 2026
Competing interests
: The authors declare no competing interests.
: This study involving human peripheral blood samples was approved by the Human Ethics Committee of Fudan University (Shanghai, China; Approval No. BE1811). Informed consent was obtained from all participants, and all methods were performed in accordance with the relevant guidelines and regulations.