Funding for this research was provided by:
National Science Foundation of China | National Natural Science Foundation of China-Yunnan Joint Fund (22025302)
National Science Foundation of China | National Natural Science Foundation of China-Yunnan Joint Fund (21873053)
Article History
Received: 25 July 2025
Revised: 27 March 2026
Accepted: 18 May 2026
First Online: 6 June 2026
Competing interests
: The authors declare no competing interests.
: All methods were carried out in accordance with relevant guidelines and regulations. This study did not involve human participants, human data, or live vertebrate animals; therefore, ethical approval and informed consent were not required.