Olli, Suvi https://orcid.org/0009-0000-0582-8729
Gustavsson, Rudolf https://orcid.org/0000-0001-8883-5958
Kivikero, Hanna https://orcid.org/0000-0002-6396-2288
Lõugas, Lembi https://orcid.org/0000-0003-2011-2141
Mannermaa, Kristiina https://orcid.org/0000-0002-8510-1120
Piličiauskienė, Giedrė https://orcid.org/0000-0002-5168-8568
Rannamäe, Eve https://orcid.org/0000-0003-1186-5512
Searle, Jeremy B. https://orcid.org/0000-0001-7710-5204
Kvist, Laura https://orcid.org/0000-0002-2108-0172
Honka, Johanna https://orcid.org/0000-0003-2536-5869
Funding for this research was provided by:
Suomen Kulttuurirahasto (00230686)
Suomen Kulttuurirahasto (00230686)
Suomen Kulttuurirahasto (00230686)
Suomen Kulttuurirahasto (00230686)
Suomen Kulttuurirahasto (00230686)
Suomen Kulttuurirahasto (00230686)
Ministry of Education and Research | Estonian Research Competency Council (TK215)
Ministry of Education and Research | Estonian Research Competency Council (TK215)
Eesti Teadusagentuur (PRG2026)
Article History
Received: 28 July 2025
Revised: 27 March 2026
Accepted: 30 March 2026
First Online: 13 April 2026
Competing interests
: The authors declare no conflict of interest.
: All analysed materials derive either from non-invasive sampling of living animals or from ancient specimens housed in collections, and were studied in accordance with relevant guidelines, permits, and institutional requirements.