Kujirai, Tomoya https://orcid.org/0000-0001-5547-9598
Kato, Junko
Yamamoto, Kyoka
Hirai, Seiya
Fujii, Takeru
Maehara, Kazumitsu https://orcid.org/0000-0002-2933-5176
Harada, Akihito https://orcid.org/0009-0003-1420-4549
Negishi, Lumi https://orcid.org/0000-0002-7297-0634
Ogasawara, Mitsuo
Yamaguchi, Yuki https://orcid.org/0000-0002-0197-614X
Ohkawa, Yasuyuki https://orcid.org/0000-0001-6440-9954
Takizawa, Yoshimasa https://orcid.org/0000-0002-6265-4830
Kurumizaka, Hitoshi https://orcid.org/0000-0001-7412-3722
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP23H05475)
MEXT | Japan Society for the Promotion of Science (JP24H02328)
MEXT | Japan Society for the Promotion of Science (JP20H03201)
MEXT | Japan Society for the Promotion of Science (JP20H05690)
MEXT | Japan Society for the Promotion of Science (JP22K15033)
MEXT | Japan Society for the Promotion of Science (JP23K17392)
MEXT | Japan Society for the Promotion of Science (JP24H00062)
MEXT | Japan Society for the Promotion of Science (JP22KJ0858)
MEXT | Japan Society for the Promotion of Science (JP22J20655)
MEXT | Japan Society for the Promotion of Science (JP22H04696)
MEXT | Japan Society for the Promotion of Science (JP23H04288)
MEXT | Japan Society for the Promotion of Science (JP21H05292)
MEXT | Japan Society for the Promotion of Science (JP23H02394)
MEXT | Japan Society for the Promotion of Science (JP20H03182)
MEXT | Japan Society for the Promotion of Science (JP23H00372)
MEXT | Japan Society for the Promotion of Science (JP24H02323)
MEXT | Japan Society for the Promotion of Science (JP22H04676)
MEXT | Japan Society for the Promotion of Science (JP22K19275)
MEXT | Japan Society for the Promotion of Science (JP22K06098)
MEXT | JST | Exploratory Research for Advanced Technology (JPMJER1901)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR24T3)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR24Q1)
Japan Agency for Medical Research and Development (JP24ama121009)
Japan Agency for Medical Research and Development (JP23gm1810008)
Japan Agency for Medical Research and Development (JP23ama121017j0001)
Japan Agency for Medical Research and Development (JP24ama121002)
MEXT | JST | Precursory Research for Embryonic Science and Technology (JPMJPR2026)
Article History
Received: 2 August 2024
Accepted: 28 April 2025
First Online: 28 May 2025
Competing interests
: The authors declare no competing interests.