Förster, Franz https://orcid.org/0000-0002-2246-6843
Emmert, David https://orcid.org/0000-0001-8149-1072
Horn, Katrin https://orcid.org/0000-0002-5307-6936
Pott, Janne
Frasnelli, Johannes
Imtiaz, Mohammed Aslam https://orcid.org/0000-0003-2762-3227
Melas, Konstantinos https://orcid.org/0009-0000-5206-5322
Talevi, Valentina
Chen, Honglei
Engel, Christoph https://orcid.org/0000-0002-7247-282X
Filosi, Michele
Fornage, Myriam https://orcid.org/0000-0003-0677-8158
Gögele, Martin https://orcid.org/0000-0003-4785-4348
Löffler, Markus
Mosley, Thomas H.
Pattaro, Cristian
Pramstaller, Peter https://orcid.org/0000-0002-9831-8302
Shrestha, Srishti
Aziz, N. Ahmad https://orcid.org/0000-0001-6184-458X
Breteler, Monique M. B. https://orcid.org/0000-0002-0626-9305
Wirkner, Kerstin
Scholz, Markus https://orcid.org/0000-0002-4059-1779
Fuchsberger, Christian https://orcid.org/0000-0002-5918-8947
Funding for this research was provided by:
Deutsche Forschungsgemeinschaft (SFB-1052/4 B11)
CoAGE graduate programme funded by the ministry for science and health of the Rhineland-Palatinate
Bundesministerium für Bildung und Forschung (01ZX1906B)
Helmholtz Association (01ER2301/14)
Wellcome Trust (225790/Z/22/Z)
RCUK | Medical Research Council (MC_UU_00002/7)
U.S. Department of Health & Human Services | National Institutes of Health (75N92022D00004)
Alzheimer's Association (AARG-19-616534)
European Research Council Starting Grant
Article History
Received: 1 October 2024
Accepted: 19 June 2025
First Online: 1 July 2025
Change Date: 11 August 2025
Change Type: Update
Change Details: In the PDF version of the article initially published, the entry for locus 9 in Table 1 appeared in bold but should have been underlined and italic (as in the online version). This has now been corrected in the PDF version of the article.
Competing interests
: M.S. received funding from Owkin for a project not related to this research. The remaining authors declare no competing interests.