Brown, Brielin C. https://orcid.org/0000-0001-5569-5223
Tokolyi, Alex https://orcid.org/0000-0003-4222-7484
Morris, John A. https://orcid.org/0000-0003-2769-8202
Lappalainen, Tuuli https://orcid.org/0000-0002-7746-8109
Knowles, David A. https://orcid.org/0000-0002-7408-146X
Funding for this research was provided by:
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (K99HG012373)
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (K99HG012792)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (R01AG057422)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (U01AG068880)
U.S. Department of Health & Human Services | NIH | National Institute of Mental Health (R01MH106842)
Article History
Received: 23 October 2023
Accepted: 15 September 2025
First Online: 31 October 2025
Competing interests
: T.L. is an advisor in Variant Bio with equity and has received speaker honoraria from AbbVie. The remaining authors declare no competing interests.