Nibe-Shirakihara, Yoichi https://orcid.org/0009-0005-9843-6674
Honda, Shinya
Arakawa, Satoko https://orcid.org/0000-0002-4375-1643
Torii, Satoru https://orcid.org/0000-0003-4841-7131
Sakurai, Hajime Tajima
Yamaguchi, Hirofumi
Oshima, Shigeru https://orcid.org/0000-0003-1186-9177
Okamoto, Ryuichi https://orcid.org/0000-0002-7047-571X
Lazarou, Michael https://orcid.org/0000-0003-2150-5545
Kawakami, Hideshi https://orcid.org/0000-0002-1405-0901
Shimizu, Shigeomi https://orcid.org/0000-0002-8437-0791
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (23H05480, 20H00467, 24K11166, 22K07345, 19K16119, 23K05748, 20K07353)
Ministry of Education, Culture, Sports, Science and Technology (22H04639, 23H04773, 24H01887)
Japan Agency for Medical Research and Development (JP23gm1410012, JP21wm0525028, JP24ama221130, JP24ek0109770)
Joint Usage/Research Program of Medical Research Institute, Institute of Science Tokyo
Department of Health | National Health and Medical Research Council (GNT1106471)
Australian Research Council Discovery Project
Article History
Received: 26 December 2023
Accepted: 11 September 2025
First Online: 20 October 2025
Competing interests
: The authors declare no competing interests.