Zhou, Peng https://orcid.org/0000-0002-3082-1462
Tsunoyama, Taka A. https://orcid.org/0000-0003-1457-7612
Kasai, Rinshi S. https://orcid.org/0000-0002-9234-3283
Hirosawa, Koichiro M. https://orcid.org/0000-0003-0170-3755
Kalay, Ziya
Aladag, Amine
Fujiwara, Takahiro K. https://orcid.org/0000-0001-5576-759X
Yokoyama, Tatsushi https://orcid.org/0000-0001-7159-9039
Sakamoto, Masayuki https://orcid.org/0000-0001-6458-113X
Kise, Ryoji https://orcid.org/0009-0000-6583-1719
Yanagawa, Masataka https://orcid.org/0000-0001-7832-0918
Inoue, Asuka https://orcid.org/0000-0003-0805-4049
Pigolotti, Simone https://orcid.org/0000-0002-6157-6906
Kusumi, Akihiro https://orcid.org/0000-0002-9558-6950
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (21K15058)
MEXT | Japan Society for the Promotion of Science (17K07333)
MEXT | Japan Society for the Promotion of Science (26870292)
MEXT | Japan Society for the Promotion of Science (20H02585)
MEXT | Japan Society for the Promotion of Science (24K01310)
MEXT | Japan Society for the Promotion of Science (18K19001)
MEXT | Japan Society for the Promotion of Science (24K18240)
MEXT | Japan Society for the Promotion of Science (JP23H02782)
MEXT | Japan Society for the Promotion of Science (24K01982)
MEXT | Japan Society for the Promotion of Science (21H04791)
MEXT | Japan Society for the Promotion of Science (24K21281)
MEXT | Japan Society for the Promotion of Science (16H06386)
MEXT | Japan Society for the Promotion of Science (22K19334)
MEXT | Japan Science and Technology Agency (JPMJAX211B)
MEXT | Japan Science and Technology Agency (JPMJAX211K)
MEXT | Japan Science and Technology Agency (JP24H00861)
MEXT | Japan Science and Technology Agency (24H01266)
MEXT | Japan Science and Technology Agency (JPMJFR215T)
MEXT | Japan Science and Technology Agency (JPMJMS2023)
Ministry of Education, Culture, Sports, Science and Technology (21H05252)
Japan Agency for Medical Research and Development (JP24wm0625119)
OIST Proof of Concept Program Technology Pioneer Fellowship: R11_61
Article History
Received: 22 August 2024
Accepted: 23 September 2025
First Online: 7 November 2025
Competing interests
: P.Z. and A.K. are inventors on a patent application (patent pending) for Xpeps and Xpep-TATs described in this study. The remaining authors declare no competing interests.