Matsumoto, Syota https://orcid.org/0000-0002-8776-2096
Takizawa, Yoshimasa https://orcid.org/0000-0002-6265-4830
Ogasawara, Mitsuo
Hashimoto, Kana
Negishi, Lumi https://orcid.org/0000-0002-7297-0634
Xu, Wenjie
Tachibana, Haruna
Yamamoto, Junpei https://orcid.org/0000-0002-5036-991X
Iwai, Shigenori
Sugasawa, Kaoru https://orcid.org/0000-0001-7937-4053
Kurumizaka, Hitoshi https://orcid.org/0000-0001-7412-3722
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP23H05475)
MEXT | Japan Society for the Promotion of Science (JP24H02328)
MEXT | Japan Society for the Promotion of Science (JP24H02319)
MEXT | Japan Society for the Promotion of Science (JP22K18034)
MEXT | Japan Society for the Promotion of Science (JP22K06098)
MEXT | JST | Exploratory Research for Advanced Technology (JPMJER1901)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR24T3)
Japan Agency for Medical Research and Development (JP24ama121009)
Japan Agency for Medical Research and Development (JP24ama121002)
MEXT | JST | Precursory Research for Embryonic Science and Technology (JPMJPR2288)
NOVARTIS Foundation (Japan) for the Promotion of Science (NOVARTIS Foundation
Sumitomo Foundation
Article History
Received: 20 March 2025
Accepted: 13 October 2025
First Online: 11 November 2025
Competing interests
: The authors declare no competing interests.