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Authors
Iwai, Hiroto https://orcid.org/0009-0008-8781-3077
Kimura, Yasuaki https://orcid.org/0000-0002-3609-602X
Honma, Masakazu
Nakamoto, Kosuke
Hashimoto, Atsushi
Motosawa, Keiichi
Atago, Takayuki
Asano, Kana
Hashiya, Fumitaka
Abe, Naoko https://orcid.org/0000-0001-6062-3515
Kobayashi, Keiko
Ogisu, Ryoko
Yamada, Hiroki https://orcid.org/0009-0009-0231-2706
Hiraishi, Keiko
Saito, Seiji https://orcid.org/0000-0002-7433-9141
Yamamoto, Junichiro
Abe, Hiroshi https://orcid.org/0000-0003-0048-3789
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Funding
Funding for this research was provided by:
Japan Agency for Medical Research and Development (JP22gm0010008)
Japan Agency for Medical Research and Development (JP22fk0310506)
Japan Agency for Medical Research and Development (JP23fk0210133)
MEXT | Japan Society for the Promotion of Science (JP22K21346)
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Article History
Received: 1 May 2024
Accepted: 23 October 2025
First Online: 19 November 2025
Competing interests
: Competing Interest Statement: H.A. is a cofounder of Crafton Biotechnology. The company focuses on the development of mRNA therapeutics. H.I., M.H., K.M., T.A., K.A., K.K., H.Y., A.H., K.H., S.S., and J.Y. are employees of Kyowa Kirin Co. Ltd. International patents application covering part of this work has been filed by Nagoya University and Kyowa Kirin Co., Ltd. The remaining authors declare no competing interests.
