Akai, Ryota https://orcid.org/0000-0002-6383-1181
Kitajima, Showa
Okubo, Kohei https://orcid.org/0009-0004-1304-5408
Serizawa, Nobuyuki https://orcid.org/0000-0002-8939-272X
Nishihara, Hirotomo https://orcid.org/0000-0003-4497-4248
Kasai, Hitoshi
Oka, Kouki https://orcid.org/0000-0001-6806-2767
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP23K17945)
MEXT | Japan Society for the Promotion of Science (JP23H03827)
MEXT | Japan Society for the Promotion of Science (JP24K01552)
MEXT | Japan Society for the Promotion of Science (JP25K21722)
MEXT | Japan Society for the Promotion of Science (JPJSBP120258801)
MEXT | Japan Society for the Promotion of Science (JP24KJ1576)
Ministry of the Environment (JPMEERF20241RA4)
MEXT | Japan Science and Technology Agency (JPMJFS2102)
Article History
Received: 14 December 2024
Accepted: 23 October 2025
First Online: 1 December 2025
Competing interests
: The authors declare no competing interests.