Kobayashi, Chisa
Ekimoto, Toru
Ooka, Koji https://orcid.org/0000-0002-0833-8853
Mifune, Junki
Matsuzawa, Kayo
Que, Lusheng
Li, Yingfang
Kusunoki, Atsuto
Morita, Takeshi
Shionoya, Kaho
Nagano, Makoto
Wakae, Kousho
Isogawa, Masanori https://orcid.org/0000-0001-8730-4303
Muramatsu, Masamichi
Ohto, Umeharu
Nomura, Norimichi https://orcid.org/0000-0002-6330-2239
Park, Sam-Yong https://orcid.org/0000-0001-6164-8896
Sureau, Camille
Arai, Munehito https://orcid.org/0000-0002-1223-2242
Ikeguchi, Mitsunori
Watashi, Koichi
Funding for this research was provided by:
Japan Agency for Medical Research and Development (JP25fk0310525)
Japan Agency for Medical Research and Development (JP25fk0310533)
Japan Agency for Medical Research and Development (JP25fk0310547)
Japan Agency for Medical Research and Development (JP23ama121007)
Japan Agency for Medical Research and Development (JP25ak0101271)
Japan Agency for Medical Research and Development (JP23fk0310517)
Japan Agency for Medical Research and Development (JP23ama121007)
Japan Agency for Medical Research and Development (JP25fk0310525)
Japan Agency for Medical Research and Development (JP25fk0310525)
Japan Agency for Medical Research and Development (JP23fk0310517)
Japan Agency for Medical Research and Development (JP23ama121023)
MEXT | Japan Society for the Promotion of Science (JP24K02290)
MEXT | Japan Society for the Promotion of Science (JP23H02724)
MEXT | Japan Society for the Promotion of Science (JP25K21767)
MEXT | Japan Society for the Promotion of Science (JP25K21782)
MEXT | Japan Society for the Promotion of Science (JP25KJ2119)
MEXT | Japan Society for the Promotion of Science (JP24K18078)
MEXT | Japan Society for the Promotion of Science (JP25H01280)
MEXT | Japan Society for the Promotion of Science (JP23H02724)
MEXT | Japan Society for the Promotion of Science (JP25K21767)
MEXT | Japan Society for the Promotion of Science (JP25H00976)
MEXT | Japan Society for the Promotion of Science (JP25H01280)
MEXT | Japan Science and Technology Agency (JPMJMI22G1)
MEXT | Japan Science and Technology Agency (JPMJSP2151)
Takeda Science Foundation (N/A)
Takeda Science Foundation (N/A)
Article History
Received: 3 June 2025
Accepted: 16 December 2025
First Online: 30 January 2026
Competing interests
: The authors declare no competing interests.