Zhang, Qiaoru
Liu, Xianhu
Guo, Yongxian https://orcid.org/0000-0001-6544-9691
Liu, Shuya
Ge, Fayuan
Gong, Yanjun https://orcid.org/0009-0005-6917-0707
Zhao, Mei
Song, Yujie
Liu, Yiwei
Sun, Di https://orcid.org/0000-0001-5966-1207
Hu, Qiongzheng https://orcid.org/0000-0003-3455-8961
Funding for this research was provided by:
National Natural Science Foundation of China (22325105)
National Natural Science Foundation of China (22106078)
Natural Science Foundation of Shandong Province (ZR2021QB031)
Natural Science Foundation of Shandong Province (ZR2022YQ12)
Taishan Scholar Foundation of Shandong Province (tsqn202306257)
Article History
Received: 30 July 2025
Accepted: 5 February 2026
First Online: 21 February 2026
Competing interests
: The authors declare no competing interests.