Calvaresi, Valeria https://orcid.org/0000-0002-1756-8853
Dello Iacono, Lucia https://orcid.org/0000-0003-2867-283X
Borghi, Sara https://orcid.org/0000-0001-5473-3240
Luzzi, Enrico
Biolchi, Alessia
Ferlenghi, Ilaria
Peschiera, Ilaria
Giusti, Fabiola
Fontana, Lucia E.
Kan, Zhong-Yuan
Spinello, Zaira
Merola, Marcello
Delany, Isabel https://orcid.org/0000-0002-9459-0595
Rand, Kasper D. https://orcid.org/0000-0002-6337-5489
Norais, Nathalie
Funding for this research was provided by:
EC | Horizon 2020 Framework Programme (675879)
Article History
Received: 17 January 2024
Accepted: 11 February 2026
First Online: 11 March 2026
Competing interests
: V.C. was an employee of the GSK group of companies and a PhD student at the University of Copenhagen at the time of this study. During this project, S.B. and I.P. held Novartis/GSK Academy Ph.D. fellowships registered at the University of Bologna (Italy); Z.S. held Novartis/GSK Academy Ph.D. fellowship registered at Sapienza, University of Rome (Italy). All other authors (except K.D.R.) are employees of the GSK group of companies. A.B., I.F., I.D. and N.N. report ownership of GSK shares and/or restricted GSK shares. All remaining authors declare no competing interests.