Ooe, Kousuke https://orcid.org/0000-0001-9632-3821
Shen, Yufan https://orcid.org/0009-0001-2097-6911
Shitara, Kazuki https://orcid.org/0000-0002-9744-8578
Maki, Rintaro
Kobayashi, Shunsuke https://orcid.org/0000-0002-4933-8817
Shimakawa, Yuichi https://orcid.org/0000-0003-1019-2512
Kan, Daisuke https://orcid.org/0000-0002-7505-0059
Etheridge, Joanne https://orcid.org/0000-0002-3199-3936
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (22KJ3209)
MEXT | Japan Society for the Promotion of Science (23K13553)
MEXT | Japan Society for the Promotion of Science (23KJ1239)
MEXT | Japan Society for the Promotion of Science (JP23H00241)
MEXT | Japan Society for the Promotion of Science (23H05457)
MEXT | Japan Society for the Promotion of Science (21H01810)
MEXT | Japan Society for the Promotion of Science (24H01190)
Ministry of Education, Culture, Sports, Science and Technology (JPMXP1020230327)
Ministry of Education, Culture, Sports, Science and Technology (JPMXP1020230325)
Ministry of Education, Culture, Sports, Science and Technology (JPMXP1122683430)
MEXT | Japan Science and Technology Agency (JPMJAP2314)
MEXT | JST | Precursory Research for Embryonic Science and Technology (JPMJPR24H3)
Department of Education and Training | Australian Research Council (FL220100202)
Article History
Received: 17 September 2025
Accepted: 26 February 2026
First Online: 30 March 2026
Competing interests
: The authors declare no competing interests.