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Yang, Mu https://orcid.org/0000-0002-6001-6827

Zhao, Rulian

Peng, Li https://orcid.org/0000-0002-7041-9267

Lv, Liting https://orcid.org/0009-0001-0928-077X

Yang, Lanyao https://orcid.org/0009-0006-5500-3892

He, Yunqi https://orcid.org/0000-0002-5265-3244

Ha, Xu https://orcid.org/0009-0007-5430-2919

Xu, Huijuan https://orcid.org/0000-0002-1858-1822

Zhang, Xiang https://orcid.org/0000-0002-1277-4615

Zhao, Peiquan https://orcid.org/0000-0002-5092-9550

Li, Shujin https://orcid.org/0000-0002-7691-3776

Yang, Zhenglin https://orcid.org/0000-0002-8656-7862

Funding for this research was provided by:

National Natural Science Foundation of China (82471102)

National Natural Science Foundation of China (82571242)

National Natural Science Foundation of China (82401292)

National Natural Science Foundation of China (82501315)

National Natural Science Foundation of China (82301231)

National Natural Science Foundation of China (82121003)

National Natural Science Foundation of China (U25A6002)

Fundamental Research Funds for the Central Universities of Ministry of Education of China

Department of Science and Technology of Sichuan Province (2023ZYD0172)

China Postdoctoral Science Foundation (2024M750365)

China Postdoctoral Science Foundation (2023M740518)

Research Fund of Sichuan Academy of Medical Sciences and Sichuan Provincial People's Hospital

National Key R&D Program of China

Jinfeng Laboratory

Text and Data Mining valid from 2026-03-24

Accepted Manuscript valid from 2026-03-24