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Ren, Junlin https://orcid.org/0000-0002-3373-6079

Wang, Sikai https://orcid.org/0000-0002-2220-0412

Zhang, Tingting

Ma, Jun https://orcid.org/0000-0003-3412-7766

Zhang, Ting https://orcid.org/0000-0001-5390-9056

Cheng, Chuyu

Zhang, Kang https://orcid.org/0000-0001-6087-2626

Xu, Changlin

van de Koppel, Johan https://orcid.org/0000-0002-0103-4275

van der Wal, Daphne https://orcid.org/0000-0002-2319-056X

Bouma, Tjeerd J.

Cheng, Fangyan

Zhuang, Ping

Zhao, Feng https://orcid.org/0000-0003-0940-6253

Funding for this research was provided by:

National Natural Science Foundation of China (32501413)

National Natural Science Foundation of China (32573518)

Natural Science Foundation of Shanghai (24ZR1481200)

Natural Science Foundation of Shanghai (23ZR1479000)

Central Public-interest Scientific Institution Basal Research Fund

the National Key R&D Program of China

Text and Data Mining valid from 2026-04-22

Accepted Manuscript valid from 2026-04-22