Pei, Xiao-Li https://orcid.org/0000-0001-5921-4466
Zhao, Pei https://orcid.org/0000-0002-8088-6141
Liu, Wen-Ting https://orcid.org/0009-0004-0037-7577
Ube, Hitoshi https://orcid.org/0000-0003-2501-7610
Lei, Zhen https://orcid.org/0000-0002-2947-108X
Ehara, Masahiro https://orcid.org/0000-0002-2185-0077
Shionoya, Mitsuhiko https://orcid.org/0000-0002-9572-4620
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP22K14691)
MEXT | Japan Society for the Promotion of Science (JP24K08443)
MEXT | Japan Society for the Promotion of Science (JP24K17663)
MEXT | Japan Society for the Promotion of Science (JP22H05133)
MEXT | Japan Society for the Promotion of Science (JP21H05022)
Ministry of Education, Culture, Sports, Science and Technology (JP16H06509)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR22B2)
Article History
Received: 9 February 2026
Accepted: 22 April 2026
First Online: 5 June 2026
Competing interests
: The authors declare no competing interests.