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Hirano, Sayuki https://orcid.org/0000-0001-7183-6604

Kondo, Yohei

Kitajima, Asayuki

Kinoshita, Noriyuki

Otani, Tetsuhisa https://orcid.org/0000-0002-2557-6722

Furuse, Mikio https://orcid.org/0000-0003-2847-8156

Ueno, Naoto https://orcid.org/0000-0002-8375-2317

Aoki, Kazuhiro https://orcid.org/0000-0001-7263-1555

Funding for this research was provided by:

MEXT | Japan Society for the Promotion of Science (JP22H02625)

MEXT | Japan Society for the Promotion of Science (JP24H01416)

MEXT | Japan Society for the Promotion of Science (25K18460)

MEXT | Japan Society for the Promotion of Science (23KJ2217)

MEXT | Japan Society for the Promotion of Science (25H01780)

MEXT | Japan Society for the Promotion of Science (21H02523)

MEXT | Japan Society for the Promotion of Science (21H02493)

Takeda Science Foundation (NA)

Takeda Science Foundation (NA)

KA was supported by grants from the International Research Collaboration Center at the National Institutes of Natural Sciences and the Nakajima Foundation.

SH was supported by grants from the International Research Collaboration Center at the National Institutes of Natural Sciences.

NU was supported by grants from the International Research Collaboration Center at the National Institutes of Natural Sciences.

Text and Data Mining valid from 2026-05-09

Accepted Manuscript valid from 2026-05-09