Watanabe, Takeo https://orcid.org/0000-0002-4562-5376
Sasaki, Yuka https://orcid.org/0000-0002-0061-0250
Zama, Takuro https://orcid.org/0000-0002-7125-2112
Matthews, Julian R. https://orcid.org/0000-0002-1895-3403
Ogawa, Daiki
Shibata, Kazuhisa https://orcid.org/0000-0002-4183-3493
Funding for this research was provided by:
National Science Foundation (BSF 2241417)
Foundation for the National Institutes of Health (R01 EY027841)
Foundation for the National Institutes of Health (R01 EY019466)
Foundation for the National Institutes of Health (R01 EY031705)
MEXT | Japan Society for the Promotion of Science (Kakenhi Grant 20H05715)
MEXT | Japan Science and Technology Agency (Moonshot R&D Grant JPMJMS2013)
Article History
Received: 23 November 2025
Accepted: 23 April 2026
First Online: 18 May 2026
Competing interests
: The authors declare no competing interests.