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Authors
Shimada, Issei S. https://orcid.org/0000-0002-8808-8419
Goto, Akari
Hashimoto, Yutaka
Inoue, Hajime
Sugawara, Takuto
Doura, Tomohiro https://orcid.org/0000-0003-1826-6010
Fujita, Tsubasa
Iwata, Toshiaki
Shimmoto, Riko
Takase, Hiroshi
Itoh, Masayuki https://orcid.org/0000-0001-7247-2259
Kiyonaka, Shigeki https://orcid.org/0000-0002-4100-6738
Kato, Yoichi https://orcid.org/0000-0002-2206-7279
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Funding
Funding for this research was provided by:
Nagoya City University (2401101)
MEXT | NINS | National Institute for Basic Biology (24NIBB460)
MEXT | Japan Society for the Promotion of Science (21K07803)
MEXT | Japan Society for the Promotion of Science (24K02425)
MEXT | Japan Society for the Promotion of Science (20K16157)
MEXT | Japan Society for the Promotion of Science (24H02265)
MEXT | Japan Society for the Promotion of Science (24H00492)
MEXT | Japan Society for the Promotion of Science (24K21823)
MEXT | Japan Society for the Promotion of Science (20K21584)
Mochida Memorial Foundation for Medical and Pharmaceutical Research
Takeda Science Foundation
the Daiko Foundation, The Hori Science and Arts Foundation, iPS Academia Japan Inc., Leave a Nest Co., Ltd. (Ikeda Rika Award) and the Nitto Foundation
Japan Agency for Medical Research and Development (24zf0127012)
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Article History
Received: 14 July 2025
Accepted: 13 May 2026
First Online: 22 May 2026
Competing interests
: I.S.S. and Y.K. submitted a patent application about dorsal brain organoid formation. The patent applicant is Nagoya City University, the inventors are Issei S. Shimada and Yoichi Kato, the application number is 2023-157480, and the status of the application is published and pending; request for examination not yet filed. The specific aspects of manuscript covered by the patent application are related to Fig. 5b–e, Supplementary Fig. , and Supplementary Fig. . The remaining authors declare no competing interests.
