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Authors
Yamori, Wataru https://orcid.org/0000-0001-7215-4736
Nakazato, Issei
Qu, Yuchen
Sanga, Yukina
Miyata, Tomoko
Uehara, Ryo
Noto, Yuma
Namba, Keiichi https://orcid.org/0000-0003-2911-5875
Fukayama, Hiroshi
Matsumura, Hiroyoshi https://orcid.org/0000-0003-0361-3796
Arimura, Shin-ichi https://orcid.org/0000-0002-9537-1626
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (24H02271)
MEXT | Japan Society for the Promotion of Science (JPJSCCA20230008)
MEXT | Japan Society for the Promotion of Science (21H02171)
MEXT | Japan Society for the Promotion of Science (24H02277)
MEXT | Japan Society for the Promotion of Science (22J20237)
MEXT | Japan Society for the Promotion of Science (25H02292, 24H02277, 24K01994, 23K18033 and 25H02292)
MEXT | JST | Advanced Low Carbon Technology Research and Development Program (JPMJAN25D2)
MEXT | JST | Program on Open Innovation Platform with Enterprises, Research Institute and Academia (JPMJOP1861)
Japan Agency for Medical Research and Development (JP21am0101117 and JP22ama121003)
Japan Agency for Medical Research and Development (JP17pc0101020)
Japan Agency for Medical Research and Development (JP23ama121001)
Osaka University (CR-22-02 and CR-23-02)
Ritsumeikan University | Ritsumeikan Global Innovation Research Organization, Ritsumeikan University
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Article History
Received: 22 October 2025
Accepted: 15 May 2026
First Online: 19 June 2026
Competing interests
: W.Y., I.N., H.F., H.M., and S.A. are applying a patent related to the finding in this paper pending in Japan (no. 2024-191646). The patent is held jointly by the University of Tokyo, Ritsumeikan University, and Kobe University. The remaining authors declare no competing interests.
