Jin, Ying
An, Hao-Jie
Zheng, Ting-Ting
Li, Jun-Jian
Gao, Jiang-Mei
Zhong, Xiu-Ling
Li, Bi-Hai
Liu, Yi-Yan
Zhuang, Xiao-Ji
Chen, Jian-Huan
Rao, Jun-Hua
Funding for this research was provided by:
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
National Natural Science Foundation of China (82471097 and 82200966)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Key Laboratory of Nonhuman Primate Research (2020B121201006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
Guangdong Provincial Science and Technology Leading Talent Project (210183503006)
GDAS' Project of Science and Technology Development (2024GDASZH-2024010101)
Article History
Received: 23 December 2025
Accepted: 11 February 2026
First Online: 20 February 2026
Competing interests
: The authors declare no competing interests.
: All authors approved the final manuscript and the submission to this journal.