Koga, Toshiki https://orcid.org/0000-0001-5728-0590
Oba, Yasuhiro https://orcid.org/0000-0002-6852-3604
Takano, Yoshinori https://orcid.org/0000-0003-1151-144X
Naraoka, Hiroshi https://orcid.org/0000-0002-2373-8759
Ogawa, Nanako O. https://orcid.org/0000-0002-3823-6444
Sasaki, Kazunori
Sato, Hajime
Yoshimura, Toshihiro https://orcid.org/0000-0003-0478-4239
Ohkouchi, Naohiko https://orcid.org/0000-0002-6355-7469
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (21J00504)
MEXT | Japan Society for the Promotion of Science (21H05414)
MEXT | Japan Society for the Promotion of Science (21KK0062)
MEXT | Japan Society for the Promotion of Science (23H00148)
Article History
Received: 8 July 2025
Accepted: 20 January 2026
First Online: 16 March 2026
Competing interests
: The authors declare no competing interests.