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Authors
Katsuda, Yousuke https://orcid.org/0000-0002-5586-4530
Kamura, Takuto
Kida, Tomoki https://orcid.org/0009-0009-4234-3160
Ohno, Rinka
Shiroto, Shuhei
Hasegawa, Yua
Utsumi, Kaito
Sakamoto, Yuki
Nakamura, Shinichiro
Nakamura, Taishi https://orcid.org/0000-0002-1205-3252
Tsujita, Kenichi
Kitamura, Yusuke
Kamiya, Yukiko https://orcid.org/0000-0001-7582-3886
Asanuma, Hiroyuki
Ihara, Toshihiro https://orcid.org/0000-0001-6236-7818
Hagihara, Masaki https://orcid.org/0000-0002-2678-825X
Sato, Shin-ichi https://orcid.org/0000-0003-3085-419X
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (20H02859, 21K18214, 23H02082, 21J11197, 23KJ1773, 20H02769, 22K05326, JP23H04067, JP21H05025)
Japan Agency for Medical Research and Development (JP20ak0101116, JP23ak0101168, JP21lm0203004, JP22ym0126806, JP22ym0126813, JP20lm0203012, JP23fk0310518, JP22fk0410055, JP23am0401007, JP23am0401007)
Naito Foundation
Terumo Foundation for Life Sciences and Arts
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Article History
Received: 10 October 2023
Accepted: 22 August 2025
First Online: 15 October 2025
Competing interests
: Y. Katsuda, Y. Kitamura, T.I., M.H. and S. Sato are coinventors on provisional patent applications (US, no. 17/287563; JP, no. 2020-552638; CN, no. 201980070388.3; EU, no. 19875072.1; HK, no. 62022046162 and PCT/JP2022/045583) filed by Kumamoto University. Y. Katsuda is a chief scientific officer and cofounder of StapleBio Inc. The other authors declare no competing interests.
