Liu, Yali https://orcid.org/0000-0001-9814-7964
Jiang, Han
Miao, Yuanxiang
Zhao, Wenli
Schneider, Ralf https://orcid.org/0000-0001-6015-7219
Yin, Liduo
Yu, Xinyue
Yu, Haiyan
Lu, Xuemei https://orcid.org/0000-0001-6044-6002
Bi, Enguang
Chen, Luonan https://orcid.org/0000-0002-3960-0068
Meyer, Axel https://orcid.org/0000-0002-0888-8193
Lin, Qiang https://orcid.org/0000-0002-9916-7761
Funding for this research was provided by:
CAS | South China Sea Institute of Oceanology, Chinese Academy of Sciences (SCSIO2024QY01)
National Natural Science Foundation of China (42270127)
National Natural Science Foundation of China (42425004, 42230409)
Article History
Received: 25 February 2025
Accepted: 24 September 2025
First Online: 11 November 2025
Competing interests
: The authors declare no competing interests.