Yang, Xiaoyu
Cai, Guanjing
Cai, Runlin
Gu, Haifeng
Chen, Yuerong
Xie, Jianmin
Hu, Zhong
Leung, Jonathan Y. S. https://orcid.org/0000-0001-5846-3401
Teeling, Hanno https://orcid.org/0000-0003-1889-7859
Wang, Hui https://orcid.org/0000-0001-7955-7239
Funding for this research was provided by:
National Natural Science Foundation of China (42476122)
National Natural Science Foundation of China (92051118)
National Natural Science Foundation of China (42476092)
National Natural Science Foundation of China (42576151)
Guangdong Science and Technology Department (2022B1515020017)
Guangdong Science and Technology Department (2025A1515010933)
Department of Education of Guangdong Province (0202202300032)
Shantou University (NTF23027T)
Deutsche Forschungsgemeinschaft (542264307)
Article History
Received: 22 April 2025
Accepted: 26 February 2026
First Online: 20 March 2026
Competing interests
: The authors declare no competing interests.