Liang, Kaiyu https://orcid.org/0000-0003-2535-8220
Zhao, Lan https://orcid.org/0009-0009-0512-3791
Zhang, Shuheng
Zheng, Luyu
Zhang, Zheyuan https://orcid.org/0009-0002-1259-5577
Wang, Shengyu
Chen, Junxin
Xu, Wenbin
Wang, Weikai
Yang, Hanshen https://orcid.org/0000-0001-5220-1996
Song, Chenxin
Qiu, Pengcheng
Zhao, Chenchen
Fang, Weifeng
Zhu, Jinjin
Fan, Shunwu https://orcid.org/0000-0003-3443-4395
Liu, Zhaoming https://orcid.org/0000-0002-7564-6207
Tang, Ruikang https://orcid.org/0000-0001-5277-7338
Zhao, Yueqi https://orcid.org/0000-0003-1867-8891
Fang, Xiangqian https://orcid.org/0000-0003-4500-8914
Funding for this research was provided by:
National Natural Science Foundation of China (U22A20282, 82472471, 22435006)
Natural Science Foundation of Zhejiang Province (LMS25B010001)
Article History
Received: 21 May 2024
Accepted: 14 August 2025
First Online: 1 October 2025
Change Date: 20 November 2025
Change Type: Update
Change Details: In the version of the article initially published, two of the files (Uncropped blots for Fig. 5k and Uncropped blots for Extended Data Fig. 1) were incorrect and have now been replaced in the HTML version of the article.
Competing interests
: The authors declare no competing interests.