Li, Zhen https://orcid.org/0000-0002-9153-3181
Cai, Rundong https://orcid.org/0009-0002-2419-4498
Wang, Xulong https://orcid.org/0000-0001-8371-0339
Shimomura, Kenji
Lu, Congwei
Yang, Zhesen https://orcid.org/0000-0002-3217-2910
Sato, Masatoshi https://orcid.org/0009-0002-3816-752X
Ma, Guancong https://orcid.org/0000-0003-3652-4351
Funding for this research was provided by:
Ministry of Science and Technology of the People’s Republic of China (2022YFA1404400)
Ministry of Science and Technology of the People’s Republic of China (2023YFA1407500)
National Natural Science Foundation of China (T2525002)
National Natural Science Foundation of China (12104450)
Research Grants Council, University Grants Committee (12300925)
Research Grants Council, University Grants Committee (JRFS2526-2S07)
Hong Kong Baptist University (RC-RSRG/23-24/SCI/01)
Hong Kong Baptist University (RC-SFCRG/23-24/R2/SCI/12)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR19T2)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR19T2)
MEXT | Japan Society for the Promotion of Science (JPMJSP2110)
MEXT | Japan Society for the Promotion of Science (JP25KJ1632)
MEXT | Japan Society for the Promotion of Science (JP24K00569)
Article History
Received: 6 May 2025
Accepted: 18 March 2026
First Online: 28 April 2026
Competing interests
: The authors declare no competing interests.