Hioki, Tomosato https://orcid.org/0000-0003-3240-7633
Tojo, Kaito https://orcid.org/0009-0005-3928-0323
Elyasi, Mehrdad https://orcid.org/0000-0002-2547-1445
Horibe, Sohei
Shimizu, Hiroki
Hoshi, Koujiro https://orcid.org/0000-0003-3817-2628
Makiuchi, Takahiko https://orcid.org/0000-0002-2155-8201
Bauer, Gerrit E. W. https://orcid.org/0000-0002-3615-8673
Saitoh, Eiji https://orcid.org/0000-0003-2675-0718
Funding for this research was provided by:
MEXT | JST | Precursory Research for Embryonic Science and Technology (JPMJPR24F9)
MEXT | Japan Society for the Promotion of Science (JP22K14584)
MEXT | Japan Society for the Promotion of Science (JP21K13847)
MEXT | Japan Society for the Promotion of Science (JP24KJ0927)
MEXT | Japan Society for the Promotion of Science (JP23KJ0607)
MEXT | Japan Society for the Promotion of Science (JP22H04965)
MEXT | Japan Society for the Promotion of Science (JP24H02231)
MEXT | Japan Society for the Promotion of Science (JP25K17943)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR20C1)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR20T2)
Article History
Received: 7 January 2026
Accepted: 14 April 2026
First Online: 1 June 2026
Competing interests
: The authors declare no competing interests.