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Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP25K00958)
MEXT | Japan Society for the Promotion of Science (24H00965)
MEXT | Japan Society for the Promotion of Science (JP23K13060)
MEXT | Japan Society for the Promotion of Science (JP23H00089)
MEXT | Japan Society for the Promotion of Science (JP22H00105)
MEXT | Japan Society for the Promotion of Science (JP25H01248)
MEXT | Japan Society for the Promotion of Science (JP24K00568)
MEXT | Japan Society for the Promotion of Science (JP23H00089)
MEXT | Japan Science and Technology Agency (JPMJPR2252)
MEXT | Japan Science and Technology Agency (JPMJCR19T5)
Article History
Received: 20 January 2026
Accepted: 21 May 2026
First Online: 17 June 2026
Competing interests
: The authors declare no competing interests.