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Authors
Freedman, Benjamin S. https://orcid.org/0000-0003-2228-7383
Bulte, Jeff W. M. https://orcid.org/0000-0003-1202-1610
Conklin, Bruce R.
Judge, Luke M.
Dwinell, Melinda R.
Geurts, Aron M.
Sitton, Madeleine J.
Mahajan, Vineet
Kiani, Samira
Gersbach, Charles A. https://orcid.org/0000-0003-1478-4013
Ebrahimkhani, Mo R.
Kelly, John J. https://orcid.org/0000-0002-3626-8629
Ronald, John A. https://orcid.org/0000-0003-3665-173X
Morizane, Ryuji
Gupta, Navin
Shakeri-Zadeh, Ali
Vo, Nicole
Saha, Krishanu https://orcid.org/0000-0003-2837-0858
Saxena, Shivani
Gamm, David M.
Sinha, Divya https://orcid.org/0000-0002-5854-0056
Tarantal, Alice F. https://orcid.org/0000-0002-8153-1094
Vandsburger, Moriel
Matsubara, Azusa
Fu, Hongxia
Tsai, Shengdar Q. https://orcid.org/0000-0001-9161-3993
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Article History
Accepted: 28 October 2025
First Online: 13 January 2026
Competing interests
: B.S.F. is an inventor on patents related to human kidney organoid differentiation and disease modelling. B.S.F. and H.F. hold ownership interest in Plurexa LLC. S.K. and M.R.E. are cofounders of GenexGen Inc and Hexembio Inc. M.R.E is a cofounder at Orgagen Biosciences Inc. R.M. is an inventor on a patent related to this work filed by President and Fellows of Harvard College and Mass General Brigham (PCT/US2018/036677 licensed to Trestle Biotherapeutics), holds a stock option in Trestle Biotherapeutics and served as a consultant to Toray Industries and Ajinomoto. K.S. is an adviser to Andson Biotechnology and Bharat Biotechnology. J.W.M.B. is a paid scientific advisory board member of Novadip Biosciences and shareholder of SuperBranche. B.R.C. is a founder and shareholder of Tenaya Therapeutics. C.A.G. is a cofounder of Tune Therapeutics, Locus Biosciences and Sollus Therapeutics, an adviser to Sarepta Therapeutics and an inventor on patent applications related to genome editing. S.Q.T. is a co-inventor on patents on GUIDE-seq, CIRCLE-seq and CHANGE-seq and is a member of the scientific advisory boards of Prime Medicine and Ensoma. D.M.G. is an inventor on a patent for production of three-dimensional retinal organoids (through the Wisconsin Alumni Research Foundation, Madison, WI (US PTO no. 9,328,328)), has an ownership interest in and receives grant support from Opsis Therapeutics LLC. The other authors declare no competing interests.
