Updates are available Correction dated 2022-04-26
Click to view Correction: https://doi.org/10.1038/s41586-021-04226-3
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Authors
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Abascal, Federico
Acosta, Reyes
Addleman, Nicholas J.
Adrian, Jessika
Afzal, Veena
Ai, Rizi
Aken, Bronwen
Akiyama, Jennifer A.
Jammal, Omar Al
Amrhein, Henry
Anderson, Stacie M.
Andrews, Gregory R.
Antoshechkin, Igor
Ardlie, Kristin G.
Armstrong, Joel
Astley, Matthew
Banerjee, Budhaditya
Barkal, Amira A.
Barnes, If H. A.
Barozzi, Iros
Barrell, Daniel
Barson, Gemma
Bates, Daniel
Baymuradov, Ulugbek K.
Bazile, Cassandra
Beer, Michael A.
Beik, Samantha
Bender, M. A.
Bennett, Ruth
Bouvrette, Louis Philip Benoit
Bernstein, Bradley E.
Berry, Andrew
Bhaskar, Anand
Bignell, Alexandra
Blue, Steven M.
Bodine, David M.
Boix, Carles
Boley, Nathan
Borrman, Tyler
Borsari, Beatrice
Boyle, Alan P.
Brandsmeier, Laurel A.
Breschi, Alessandra
Bresnick, Emery H.
Brooks, Jason A.
Buckley, Michael
Burge, Christopher B.
Byron, Rachel
Cahill, Eileen
Cai, Lingling
Cao, Lulu
Carty, Mark
Castanon, Rosa G.
Castillo, Andres
Chaib, Hassan
Chan, Esther T.
Chee, Daniel R.
Chee, Sora
Chen, Hao
Chen, Huaming
Chen, Jia-Yu
Chen, Songjie
Cherry, J. Michael
Chhetri, Surya B.
Choudhary, Jyoti S.
Chrast, Jacqueline
Chung, Dongjun
Clarke, Declan
Cody, Neal A. L.
Coppola, Candice J.
Coursen, Julie
D’Ippolito, Anthony M.
Dalton, Stephen
Danyko, Cassidy
Davidson, Claire
Davila-Velderrain, Jose
Davis, Carrie A.
Dekker, Job
Deran, Alden
DeSalvo, Gilberto
Despacio-Reyes, Gloria
Dewey, Colin N.
Dickel, Diane E.
Diegel, Morgan
Diekhans, Mark
Dileep, Vishnu
Ding, Bo
Djebali, Sarah
Dobin, Alexander
Dominguez, Daniel
Donaldson, Sarah
Drenkow, Jorg
Dreszer, Timothy R.
Drier, Yotam
Duff, Michael O.
Dunn, Douglass
Eastman, Catharine
Ecker, Joseph R.
Edwards, Matthew D.
El-Ali, Nicole
Elhajjajy, Shaimae I.
Elkins, Keri
Emili, Andrew
Epstein, Charles B.
Evans, Rachel C.
Ezkurdia, Iakes
Fan, Kaili
Farnham, Peggy J.
Farrell, Nina P.
Feingold, Elise A.
Ferreira, Anne-Maud
Fisher-Aylor, Katherine
Fitzgerald, Stephen
Flicek, Paul
Foo, Chuan Sheng
Fortier, Kevin
Frankish, Adam
Freese, Peter
Fu, Shaliu
Fu, Xiang-Dong
Fu, Yu
Fukuda-Yuzawa, Yoko
Fulciniti, Mariateresa
Funnell, Alister P. W.
Gabdank, Idan
Galeev, Timur
Gao, Mingshi
Giron, Carlos Garcia
Garvin, Tyler H.
Gelboin-Burkhart, Chelsea Anne
Georgolopoulos, Grigorios
Gerstein, Mark B.
Giardine, Belinda M.
Gifford, David K.
Gilbert, David M.
Gilchrist, Daniel A.
Gillespie, Shawn
Gingeras, Thomas R.
Gong, Peng
Gonzalez, Alvaro
Gonzalez, Jose M.
Good, Peter
Goren, Alon
Gorkin, David U.
Graveley, Brenton R.
Gray, Michael
Greenblatt, Jack F.
Griffiths, Ed
Groudine, Mark T.
Grubert, Fabian
Gu, Mengting
Guigó, Roderic
Guo, Hongbo
Guo, Yu
Guo, Yuchun
Gursoy, Gamze
Gutierrez-Arcelus, Maria
Halow, Jessica
Hardison, Ross C.
Hardy, Matthew
Hariharan, Manoj
Harmanci, Arif
Harrington, Anne
Harrow, Jennifer L.
Hashimoto, Tatsunori B.
Hasz, Richard D.
Hatan, Meital
Haugen, Eric
Hayes, James E.
He, Peng
He, Yupeng
Heidari, Nastaran
Hendrickson, David
Heuston, Elisabeth F.
Hilton, Jason A.
Hitz, Benjamin C.
Hochman, Abigail
Holgren, Cory
Hou, Lei
Hou, Shuyu
Hsiao, Yun-Hua E.
Hsu, Shanna
Huang, Hui
Hubbard, Tim J.
Huey, Jack
Hughes, Timothy R.
Hunt, Toby
Ibarrientos, Sean
Issner, Robbyn
Iwata, Mineo
Izuogu, Osagie
Jaakkola, Tommi
Jameel, Nader
Jansen, Camden
Jiang, Lixia
Jiang, Peng
Johnson, Audra
Johnson, Rory
Jungreis, Irwin
Kadaba, Madhura
Kasowski, Maya
Kasparian, Mary
Kato, Momoe
Kaul, Rajinder
Kawli, Trupti
Kay, Michael
Keen, Judith C.
Keles, Sunduz
Keller, Cheryl A.
Kelley, David
Kellis, Manolis
Kheradpour, Pouya
Kim, Daniel Sunwook
Kirilusha, Anthony
Klein, Robert J.
Knoechel, Birgit
Kuan, Samantha
Kulik, Michael J.
Kumar, Sushant
Kundaje, Anshul
Kutyavin, Tanya
Lagarde, Julien
Lajoie, Bryan R.
Lambert, Nicole J.
Lazar, John
Lee, Ah Young
Lee, Donghoon
Lee, Elizabeth
Lee, Jin Wook
Lee, Kristen
Leslie, Christina S.
Levy, Shawn
Li, Bin
Li, Hairi
Li, Nan
Li, Shantao
Li, Xiangrui
Li, Yang I.
Li, Ying
Li, Yining
Li, Yue
Lian, Jin
Libbrecht, Maxwell W.
Lin, Shin
Lin, Yiing
Liu, Dianbo
Liu, Jason
Liu, Peng
Liu, Tingting
Liu, X. Shirley
Liu, Yan
Liu, Yaping
Long, Maria
Lou, Shaoke
Loveland, Jane
Lu, Aiping
Lu, Yuheng
Lécuyer, Eric
Ma, Lijia
Mackiewicz, Mark
Mannion, Brandon J.
Mannstadt, Michael
Manthravadi, Deepa
Marinov, Georgi K.
Martin, Fergal J.
Mattei, Eugenio
McCue, Kenneth
McEown, Megan
McVicker, Graham
Meadows, Sarah K.
Meissner, Alex
Mendenhall, Eric M.
Messer, Christopher L.
Meuleman, Wouter
Meyer, Clifford
Miller, Steve
Milton, Matthew G.
Mishra, Tejaswini
Moore, Dianna E.
Moore, Helen M.
Moore, Jill E.
Moore, Samuel H.
Moran, Jennifer
Mortazavi, Ali
Mudge, Jonathan M.
Munshi, Nikhil
Murad, Rabi
Myers, Richard M.
Nandakumar, Vivek
Nandi, Preetha
Narasimha, Anil M.
Narayanan, Aditi K.
Naughton, Hannah
Navarro, Fabio C. P.
Navas, Patrick
Nazarovs, Jurijs
Nelson, Jemma
Neph, Shane
Neri, Fidencio Jun
Nery, Joseph R.
Nesmith, Amy R.
Newberry, J. Scott
Newberry, Kimberly M.
Ngo, Vu
Nguyen, Rosy
Nguyen, Thai B.
Nguyen, Tung
Nishida, Andrew
Noble, William S.
Novak, Catherine S.
Novoa, Eva Maria
Nuñez, Briana
O’Donnell, Charles W.
Olson, Sara
Onate, Kathrina C.
Otterman, Ericka
Ozadam, Hakan
Pagan, Michael
Palden, Tsultrim
Pan, Xinghua
Park, Yongjin
Partridge, E. Christopher
Paten, Benedict
Pauli-Behn, Florencia
Pazin, Michael J.
Pei, Baikang
Pennacchio, Len A.
Perez, Alexander R.
Perry, Emily H.
Pervouchine, Dmitri D.
Phalke, Nishigandha N.
Pham, Quan
Phanstiel, Doug H.
Plajzer-Frick, Ingrid
Pratt, Gabriel A.
Pratt, Henry E.
Preissl, Sebastian
Pritchard, Jonathan K.
Pritykin, Yuri
Purcaro, Michael J.
Qin, Qian
Quinones-Valdez, Giovanni
Rabano, Ines
Radovani, Ernest
Raj, Anil
Rajagopal, Nisha
Ram, Oren
Ramirez, Lucia
Ramirez, Ricardo N.
Rausch, Dylan
Raychaudhuri, Soumya
Raymond, Joseph
Razavi, Rozita
Reddy, Timothy E.
Reimonn, Thomas M.
Ren, Bing
Reymond, Alexandre
Reynolds, Alex
Rhie, Suhn K.
Rinn, John
Rivera, Miguel
Rivera-Mulia, Juan Carlos
Roberts, Brian S.
Rodriguez, Jose Manuel
Rozowsky, Joel
Ryan, Russell
Rynes, Eric
Salins, Denis N.
Sandstrom, Richard
Sasaki, Takayo
Sathe, Shashank
Savic, Daniel
Scavelli, Alexandra
Scheiman, Jonathan
Schlaffner, Christoph
Schloss, Jeffery A.
Schmitges, Frank W.
See, Lei Hoon
Sethi, Anurag
Setty, Manu
Shafer, Anthony
Shan, Shuo
Sharon, Eilon
Shen, Quan
Shen, Yin
Sherwood, Richard I.
Shi, Minyi
Shin, Sunyoung
Shoresh, Noam
Siebenthall, Kyle
Sisu, Cristina
Slifer, Teri
Sloan, Cricket A.
Smith, Anna
Snetkova, Valentina
Snyder, Michael P.
Spacek, Damek V.
Srinivasan, Sharanya
Srivas, Rohith
Stamatoyannopoulos, George
Stamatoyannopoulos, John A.
Stanton, Rebecca
Steffan, Dave
Stehling-Sun, Sandra
Strattan, J. Seth
Su, Amanda
Sundararaman, Balaji
Suner, Marie-Marthe
Syed, Tahin
Szynkarek, Matt
Tanaka, Forrest Y.
Tenen, Danielle
Teng, Mingxiang
Thomas, Jeffrey A.
Toffey, Dave
Tress, Michael L.
Trout, Diane E.
Trynka, Gosia
Tsuji, Junko
Upchurch, Sean A.
Ursu, Oana
Uszczynska-Ratajczak, Barbara
Uziel, Mia C.
Valencia, Alfonso
Biber, Benjamin Van
van der Velde, Arjan G.
Van Nostrand, Eric L.
Vaydylevich, Yekaterina
Vazquez, Jesus
Victorsen, Alec
Vielmetter, Jost
Vierstra, Jeff
Visel, Axel
Vlasova, Anna
Vockley, Christopher M.
Volpi, Simona
Vong, Shinny
Wang, Hao
Wang, Mengchi
Wang, Qin
Wang, Ruth
Wang, Tao
Wang, Wei
Wang, Xiaofeng
Wang, Yanli
Watson, Nathaniel K.
Wei, Xintao
Wei, Zhijie
Weisser, Hendrik
Weissman, Sherman M.
Welch, Rene
Welikson, Robert E.
Weng, Zhiping
Westra, Harm-Jan
Whitaker, John W.
White, Collin
White, Kevin P.
Wildberg, Andre
Williams, Brian A.
Wine, David
Witt, Heather N.
Wold, Barbara
Wolf, Maxim
Wright, James
Xiao, Rui
Xiao, Xinshu
Xu, Jie
Xu, Jinrui
Yan, Koon-Kiu
Yan, Yongqi
Yang, Hongbo
Yang, Xinqiong
Yang, Yi-Wen
Yardımcı, Galip Gürkan
Yee, Brian A.
Yeo, Gene W.
Young, Taylor
Yu, Tianxiong
Yue, Feng
Zaleski, Chris
Zang, Chongzhi
Zeng, Haoyang
Zeng, Weihua
Zerbino, Daniel R.
Zhai, Jie
Zhan, Lijun
Zhan, Ye
Zhang, Bo
Zhang, Jialing
Zhang, Jing
Zhang, Kai
Zhang, Lijun
Zhang, Peng
Zhang, Qi
Zhang, Xiao-Ou
Zhang, Yanxiao
Zhang, Zhizhuo
Zhao, Yuan
Zheng, Ye
Zhong, Guoqing
Zhou, Xiao-Qiao
Zhu, Yun
Zimmerman, Jared
Moore, Jill E. https://orcid.org/0000-0002-3023-0806
Purcaro, Michael J. https://orcid.org/0000-0002-4735-4215
Pratt, Henry E.
Epstein, Charles B. https://orcid.org/0000-0001-8358-8345
Shoresh, Noam
Adrian, Jessika
Kawli, Trupti
Davis, Carrie A.
Dobin, Alexander
Kaul, Rajinder https://orcid.org/0000-0002-7895-9284
Halow, Jessica
Van Nostrand, Eric L.
Freese, Peter
Gorkin, David U. https://orcid.org/0000-0003-4944-4107
Shen, Yin https://orcid.org/0000-0001-9901-5613
He, Yupeng
Mackiewicz, Mark https://orcid.org/0000-0002-7088-1902
Pauli-Behn, Florencia
Williams, Brian A.
Mortazavi, Ali https://orcid.org/0000-0002-4259-6362
Keller, Cheryl A. https://orcid.org/0000-0001-6594-0245
Zhang, Xiao-Ou
Elhajjajy, Shaimae I.
Huey, Jack
Dickel, Diane E. https://orcid.org/0000-0001-5497-6824
Snetkova, Valentina
Wei, Xintao
Wang, Xiaofeng
Rivera-Mulia, Juan Carlos https://orcid.org/0000-0002-7566-3875
Rozowsky, Joel https://orcid.org/0000-0002-3565-0762
Zhang, Jing
Chhetri, Surya B.
Zhang, Jialing
Victorsen, Alec
White, Kevin P.
Visel, Axel https://orcid.org/0000-0002-4130-7784
Yeo, Gene W. https://orcid.org/0000-0002-0799-6037
Burge, Christopher B. https://orcid.org/0000-0001-9047-5648
Lécuyer, Eric
Gilbert, David M. https://orcid.org/0000-0001-8087-9737
Dekker, Job https://orcid.org/0000-0001-5631-0698
Rinn, John https://orcid.org/0000-0002-7231-7539
Mendenhall, Eric M. https://orcid.org/0000-0002-7395-6295
Ecker, Joseph R. https://orcid.org/0000-0001-5799-5895
Kellis, Manolis https://orcid.org/0000-0001-7113-9630
Klein, Robert J. https://orcid.org/0000-0003-3539-5391
Noble, William S.
Kundaje, Anshul https://orcid.org/0000-0003-3084-2287
Guigó, Roderic
Farnham, Peggy J. https://orcid.org/0000-0003-4469-7914
Cherry, J. Michael https://orcid.org/0000-0001-9163-5180
Myers, Richard M.
Ren, Bing https://orcid.org/0000-0002-5435-1127
Graveley, Brenton R.
Gerstein, Mark B.
Pennacchio, Len A. https://orcid.org/0000-0002-8748-3732
Snyder, Michael P. https://orcid.org/0000-0003-0784-7987
Bernstein, Bradley E. https://orcid.org/0000-0002-5726-6278
Wold, Barbara
Hardison, Ross C. https://orcid.org/0000-0003-4084-7516
Gingeras, Thomas R.
Stamatoyannopoulos, John A.
Weng, Zhiping https://orcid.org/0000-0002-3032-7966
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Article History
Received: 26 August 2017
Accepted: 27 May 2020
First Online: 29 July 2020
Change Date: 27 August 2020
Change Type: Update
Change Details: The Supplementary Information file initially published was corrupted and replaced online.
Change Date: 26 April 2022
Change Type: Correction
Change Details: A Correction to this paper has been published:
Change Details: https://doi.org/10.1038/s41586-021-04226-3
Competing interests
: B.E.B. declares outside interests in Fulcrum Therapeutics, 1CellBio, HiFiBio, Arsenal Biosciences, Cell Signaling Technologies, BioMillenia, and Nohla Therapeutics. P. Flicek is a member of the Scientific Advisory Boards of Fabric Genomics, Inc. and Eagle Genomics, Ltd. M.P.S. is cofounder of Personalis, SensOmics, Mirvie, Qbio, January, Filtircine, and Genome Heart. He serves on the scientific advisory board of these companies and Genapsys and Jupiter. Z. Weng is a cofounder of Rgenta Therapeutics and she serves on its scientific advisory board. G.W.Y. is co-founder, member of the Board of Directors, on the SAB, equity holder, and paid consultant for Locana and Eclipse BioInnovations, and a visiting professor at the National University of Singapore. G.W.Y.’s interests have been reviewed and approved by the University of California, San Diego in accordance with its conflict of interest policies. E.L.V.N. is co-founder, member of the Board of Directors, on the SAB, equity holder, and paid consultant for Eclipse BioInnovations. E.L.V.N.’s interests have been reviewed and approved by the University of California, San Diego in accordance with its conflict of interest policies. B.R. is a co-founder and member of SAB of Arima Genomics, Inc. The authors declare no other competing financial interests.
