Ochi, Yotaro https://orcid.org/0000-0001-8472-6164
Liew-Littorin, Markus https://orcid.org/0009-0007-5271-1321
Nannya, Yasuhito
Bengtzen, Sofia
Piauger, Benedicte
Deneberg, Stefan
Jädersten, Martin
Lazarevic, Vladimir https://orcid.org/0000-0002-1782-4423
Cammenga, Jörg
Robelius, Anna https://orcid.org/0000-0002-8853-1863
Wennström, Lovisa https://orcid.org/0009-0002-7300-5438
Ölander, Emma
Kasahara, Senji
Hiramoto, Nobuhiro https://orcid.org/0000-0002-0472-6680
Kanemura, Nobuhiro https://orcid.org/0000-0001-8449-9918
Sezaki, Nobuo
Sakurada, Maki
Iwasaki, Makoto https://orcid.org/0000-0002-7564-157X
Kanda, Junya https://orcid.org/0000-0002-6704-3633
Ueda, Yasunori
Yoshihara, Satoshi https://orcid.org/0000-0002-8537-2422
Erkers, Tom https://orcid.org/0000-0002-2754-6287
Struyf, Nona https://orcid.org/0000-0002-6975-0753
Watanabe, Yu https://orcid.org/0009-0001-5957-440X
Motomura, Masanori
Nakagawa, Masahiro M. https://orcid.org/0000-0002-6420-2555
Saiki, Ryunosuke https://orcid.org/0000-0001-6984-7187
Fukushima, Hidehito
Okazaki, Koji
Morimoto, Suguru
Yoda, Akinori
Okuda, Rurika https://orcid.org/0000-0002-7124-4171
Komatsu, Shintaro
Xie, Guoxiang https://orcid.org/0009-0003-4240-3625
Österroos, Albin https://orcid.org/0000-0001-8749-7299
Kon, Ayana
Zhao, Lanying
Shiraishi, Yuichi https://orcid.org/0000-0001-6144-5845
Ishikawa, Takayuki
Miyano, Satoru https://orcid.org/0000-0002-1753-6616
Katayama, Kotoe https://orcid.org/0000-0002-3966-1231
Imoto, Seiya
Matsuda, Shuichi https://orcid.org/0000-0003-0802-1255
Takaori-Kondo, Akifumi https://orcid.org/0000-0001-7678-4284
Aburatani, Hiroyuki
Suzuki, Hiroshi I. https://orcid.org/0000-0003-4682-5086
Kallioniemi, Olli
Juliusson, Gunnar
Höglund, Martin https://orcid.org/0000-0003-2468-0226
Lehmann, Sören https://orcid.org/0009-0009-3243-3947
Ogawa, Seishi https://orcid.org/0000-0002-7778-5374
Article History
Received: 28 February 2025
Accepted: 22 May 2026
First Online: 8 July 2026
Ethics
Competing interests: Y.O. has received research funding from Nippon Shinyaku. Y.N. has held leadership positions or advisory roles for Otsuka Pharmaceutical and Bristol Myers Squibb, and has received honoraria from Bristol Myers Squibb, Takeda Pharmaceuticals, Daiichi Sankyo, Kyowa Kirin, Novartis, Pfizer, Nippon Shinyaku, Otsuka Pharmaceutical, AstraZeneca, Astellas Pharma and AbbVie. S. Kasahara has received honoraria from Daiichi Sankyo and AbbVie and funding for clinical trials from Astellas, Novartis and Incyte Biosciences Japan. J.K. has received honoraria from Daiichi Sankyo and Asahi Kasei Pharma. S.Y. has received honoraria from Bristol Myers Squibb, Janssen Pharmaceutical and Novartis. A.Y. was employed in an endowed department funded by Chordia Therapeutics. H.A. has served in an advisory role for and has received research funding from Konica Minolta, and has an endowed chair funded by Chugai Pharmaceutical. H.I.S. has received research funding from FUJIFILM. S.O. has received research funding from Chordia Therapeutics, Eisai and Otsuka Pharmaceutical, consulting fees from Chordia Therapeutics, Eisai and Montage Bio and owns stock in Asahi Genomics. The aforementioned companies were not involved in any aspect of this study. A patent application related to this work has been filed by Kyoto University. The remaining authors declare no competing interests.