Swanson, Elliott G. https://orcid.org/0000-0002-0351-6446
Mao, Yizi https://orcid.org/0000-0001-5743-6831
Mallory, Benjamin J. https://orcid.org/0000-0003-1572-6181
Vollger, Mitchell R. https://orcid.org/0000-0002-8651-1615
Bohaczuk, Stephanie C. https://orcid.org/0000-0001-8802-6579
Oliveira, Christopher B. https://orcid.org/0000-0003-0834-3625
Lyon, Daniel B.
Ranchalis, Jane
Parmalee, Nancy L.
Cohen, Barak A. https://orcid.org/0000-0002-3350-2715
Bennett, James T. https://orcid.org/0000-0003-2843-5594
Stergachis, Andrew B. https://orcid.org/0000-0002-1299-3674
Funding for this research was provided by:
U.S. Department of Health & Human Services | NIH | NIH Office of the Director (UM1DA058220)
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (1DP5OD029630)
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (T32HG000035)
U.S. Department of Health & Human Services | NIH | National Institute on Aging (P30AG066509)
Burroughs Wellcome Fund
Pew Charitable Trusts
Shurl and Kay Curci Foundation
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (2T32GM007454)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (1K99GM155552)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (2T32GM007454)
Article History
Received: 4 September 2025
Accepted: 20 October 2025
First Online: 3 December 2025
Competing interests
: A.B.S., E.G.S. and Y.M. are co-inventors on the US Provisional Patent Application 63/687,924 which includes discoveries described in this manuscript regarding ‘Chromatin Stenciling’™. The other authors declare no competing interests.