Sakaue, Saori http://orcid.org/0000-0003-3618-9717
Kanai, Masahiro http://orcid.org/0000-0001-5165-4408
Tanigawa, Yosuke http://orcid.org/0000-0001-9759-157X
Karjalainen, Juha
Kurki, Mitja
Koshiba, Seizo
Narita, Akira http://orcid.org/0000-0002-9076-0710
Konuma, Takahiro
Yamamoto, Kenichi
Akiyama, Masato
Ishigaki, Kazuyoshi http://orcid.org/0000-0003-2881-0657
Suzuki, Akari
Suzuki, Ken http://orcid.org/0000-0003-1065-3593
Obara, Wataru
Yamaji, Ken
Takahashi, Kazuhisa
Asai, Satoshi
Takahashi, Yasuo http://orcid.org/0000-0003-2543-7543
Suzuki, Takao
Shinozaki, Nobuaki
Yamaguchi, Hiroki http://orcid.org/0000-0002-0571-7495
Minami, Shiro
Murayama, Shigeo
Yoshimori, Kozo
Nagayama, Satoshi
Obata, Daisuke
Higashiyama, Masahiko
Masumoto, Akihide
Koretsune, Yukihiro
,
Ito, Kaoru http://orcid.org/0000-0003-1843-773X
Terao, Chikashi http://orcid.org/0000-0002-6452-4095
Yamauchi, Toshimasa
Komuro, Issei http://orcid.org/0000-0002-0714-7182
Kadowaki, Takashi
Tamiya, Gen
Yamamoto, Masayuki http://orcid.org/0000-0002-9073-9436
Nakamura, Yusuke
Kubo, Michiaki http://orcid.org/0000-0002-0095-2322
Murakami, Yoshinori http://orcid.org/0000-0002-2826-4396
Yamamoto, Kazuhiko
Kamatani, Yoichiro
Palotie, Aarno http://orcid.org/0000-0002-2527-5874
Rivas, Manuel A. http://orcid.org/0000-0003-1457-9925
Daly, Mark J. http://orcid.org/0000-0002-0949-8752
Matsuda, Koichi http://orcid.org/0000-0001-7292-2686
Okada, Yukinori http://orcid.org/0000-0002-0311-8472
Funding for this research was provided by:
Japan Society for the Promotion of Science London (19H01021, 20K21834)
Japan Agency for Medical Research and Development (JP20km0405211, JP20ek0109413, JP20ek0410075, JP20gm4010006, and JP20km0405217)
Takeda Science Foundation
Bioinformatics Initiative of Osaka University Graduate School of Medicine, Osaka University
Mochida Memorial Foundation for Medical and Pharmaceutical Research
Astellas Foundation for Research on Metabolic Disorders
Kanae Foundation for the Promotion of Medical Science
The JCR Grant for Promoting Basic Rheumatology
Heiwa Nakajima Foundation
the Masason Foundation
Funai Overseas Scholarship from the Funai Foundation for Information Technology
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (R01HG010140)
a National Institute of Health center for Multi- and Cross-population Mapping of Mendelian and Complex Diseases grant
Article History
Received: 20 October 2020
Accepted: 4 August 2021
First Online: 30 September 2021
Competing interests
: M.A.R. is on the SAB of 54Gene and the Computational Advisory Board for Goldfinch Bio and has advised BioMarin, Third Rock Ventures, MazeTx and Related Sciences. The funders had no role in study design, data collection and analysis, decision to publish or preparation of the manuscript. The remaining authors declare no competing interests.