Fu, Boyang https://orcid.org/0000-0002-0082-8735
Pazokitoroudi, Ali https://orcid.org/0000-0002-2839-2291
Shi, Zhuozheng https://orcid.org/0000-0002-9769-9027
Kar, Asha https://orcid.org/0009-0000-1717-1744
Xue, Albert
Anand, Aakarsh https://orcid.org/0009-0008-9581-3827
Anand, Prateek https://orcid.org/0009-0004-8455-1308
Liu, Zhengtong
Border, Richard https://orcid.org/0000-0002-6293-2968
Pajukanta, Päivi https://orcid.org/0000-0002-6423-8056
Zaitlen, Noah
Sankararaman, Sriram https://orcid.org/0000-0003-1586-9641
Funding for this research was provided by:
National Science Foundation (CAREER-1943497)
National Science Foundation (CAREER-1943497)
National Science Foundation (CAREER-1943497)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM125055)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM153406)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM125055)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM125055)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM153406)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM153406)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (R01GM142112)
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (GM153406)
U.S. Department of Health & Human Services | NIH | National Human Genome Research Institute (HG006399)
UCLA DYA.
U.S. Department of Health & Human Services | NIH | National Institute of Mental Health (R01MH122688)
U.S. Department of Health & Human Services | NIH | National Institute of Mental Health (U01MH126798)
U.S. Department of Health & Human Services | NIH | National Heart, Lung, and Blood Institute (R01HL170604)
U.S. Department of Health & Human Services | NIH | National Institute of Diabetes and Digestive and Kidney Diseases (R01DK132775)
Article History
Received: 10 September 2023
Accepted: 16 October 2025
First Online: 9 December 2025
Competing interests
: The authors declare no competing interests.