Tajima, Masaki https://orcid.org/0000-0003-3407-0229
Hao, He
Zhang, Baihao https://orcid.org/0000-0002-3456-2509
Matsuoka, Yuta https://orcid.org/0000-0001-9860-3851
Sonomura, Kazuhiro
Imami, Koshi https://orcid.org/0000-0002-7451-4982
Isobe, Yosuke
Maeda, Rae
Lin, Yu-Hsien https://orcid.org/0000-0002-5303-1084
Shimba, Akihiro
Kato, Ryoma
Costa Cruz, Pedro Henrique
Washizu, Sayaka
Ikejiri, Yosuke
Morinibu, Akiyo
Barker, Clive Steven
Seita, Jun https://orcid.org/0000-0002-3008-3615
Wu, Yibo
Ito, Satomi
Narushima, Seiko
Nakano, Rei https://orcid.org/0000-0003-0255-6347
Maruya, Mikako
Kobayashi, Wakana
Narayanan, Sai Shanmukha Priya https://orcid.org/0009-0006-8774-6746
Shaheen, Jumana https://orcid.org/0009-0006-8997-7337
Neyama, Hiroyuki
Yamada, Ken-ichi https://orcid.org/0000-0003-2100-8477
Arita, Makoto https://orcid.org/0000-0001-9902-0463
Sugiura, Yuki
Fagarasan, Sidonia https://orcid.org/0000-0003-2086-6402
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (24K10092)
MEXT | Japan Society for the Promotion of Science (19K16881)
MEXT | Japan Society for the Promotion of Science (22K15501)
MEXT | Japan Society for the Promotion of Science (25H01031)
Mochida Memorial Foundation for Medical and Pharmaceutical Research
Astellas Foundation for Research on Metabolic Disorders
MEXT | RIKEN
Takeda Medical Research Foundation
CCII fellowship
MEXT | JST | Exploratory Research for Advanced Technology (JPMJER2101)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR24T6)
Japan Agency for Medical Research and Development (JP25gm2010001)
Japan Agency for Medical Research and Development (25zf0127003)
Japan Agency for Medical Research and Development (25zf0127007)
Yanai Fund
MEXT | Japan Science and Technology Agency (JPMJSP2110)
Article History
Received: 18 August 2025
Accepted: 12 March 2026
First Online: 13 April 2026
Competing interests
: The authors declare no competing interests.