Kim, Sangil https://orcid.org/0000-0003-2758-8943
Farrell, Brian D. https://orcid.org/0000-0002-6843-0539
Funding for this research was provided by:
Harvard University | Faculty of Arts and Sciences (OEB Graduate Research Fund)
Harvard University | Faculty of Arts and Sciences (Dean’s Competitive Fund for Promising Scholarship)
Harvard University | Faculty of Arts and Sciences (Dean’s Competitive Fund for Promising Scholarship)
National Research Foundation of Korea (2019R1A6A1A10073437)
MOE | National Institute of Biological Resources (NIBR202405101)
Harvard University | Museum of Comparative Zoology, Harvard University (Putnam Expedition Grant)
Harvard University | Museum of Comparative Zoology, Harvard University (Wetmore Colles Fund)
Harvard University | Museum of Comparative Zoology, Harvard University (Putnam Expedition Grant)
Harvard University | Museum of Comparative Zoology, Harvard University (Wetmore Colles Fund)
Article History
Received: 18 July 2025
Accepted: 3 November 2025
First Online: 10 December 2025
Competing interests
: The authors declare no competing interests.